Two 11-week-old white tiger cubs that died at Lahore Zoo last month appear to have died of Covid-19, officials said.

The cubs died in the zoo on Jan 30, four days after beginning treatment for what officials thought was feline panleukopenia virus, a disease that zoo officials said is common in Pakistan and targets cats' immune system.

But an autopsy found the cubs' lungs were badly damaged and they were suffering from severe infection, with pathologists concluding they died from Covid-19.

Although no PCR test for the novel coronavirus was conducted, zoo deputy director Kiran Saleem told Reuters the zoo believes the cubs were the victims of the pandemic that has killed 12,276 people in Pakistan.

“After their death, the zoo administration conducted tests of all officials, and six were tested positive, including one official who handled the cubs,” Saleem said.

“It strengthens the findings of the autopsy. The cubs probably caught the virus from the person handling and feeding them.”

Pakistan's zoos regularly draw the ire of animal rights activists, who say hundreds of animals have died from poor living conditions there.

“The last two white tiger cubs have died at Lahore zoo and once again the negligence of the management and authorities has come out,” Zufishan Anushay, founder of JFK (Justice for Kiki) Animal Rescue And Shelter, told Reuters.

“White tigers are extremely rare and need a specific habitat and environment to live a healthy life. By caging them in unhygienic conditions with no medical arrangements, we will keep witnessing these incidents," she said.

"Covid-19 is a new virus, and the world is making policies for humans," she said. "It should not forget animals in pet shops, zoos and everywhere else."

Saleem rejected the allegations of neglect at the zoo, telling Reuters that animal rights activists were welcome to visit and check the facility's safety and hygiene protocols themselves.

In December, two Himalayan brown bears were airlifted out of the Islamabad Zoo to a sanctuary in Jordan.

That rescue came weeks after an elephant Kaavan was moved to a sanctuary in Cambodia, the culmination of a years-long campaign that included US pop star Cher.

At the Peshawar Zoo, officials have said four giraffes died in 2020. Last year, two lions at Islamabad Zoo suffocated when workers lit fires in their cages.