Syrian government forces and their allies regained control on Saturday of a predominantly Christian central town that sleeper cells of the militant Islamic State (IS) group captured late last month.

The government-run Syrian Central Military Media said “the Syrian army and its allies have restored security and stability to Qaryatayn” after clearing the town of out IS fighters.

The capture of Qaryatayn came after nearly three weeks of fighting that saw IS capture areas it had earlier lost in an offensive by Syrian government forces and Iranian-backed militiamen under the cover of Russian airstrikes.

Earlier this week, IS lost control of the northern city of Raqqa, the de facto capital of its self-declared caliphate, after a four-month offensive by the United States (US)-backed Kurdish-led Syrian Democratic Forces. Last week, IS lost its stronghold of Mayadeen in eastern Syria.

IS still holds parts of Syria's eastern Deir el-Zour province and Iraq's Anbar province, as well as small, scattered pockets elsewhere. The loss of Qaryatayn is another blow for the extremist group.

Earlier this month, the Russian military accused the US of helping IS launch a series of attacks against Syrian troops. Moscow said that IS launched a series of attacks in late September week from the area around Tanf near Syria's border with Jordan, where US military advisers are based.

The Russians said the IS attacks near Qaryatayn in the Homs province, and a key highway linking Palmyra and Deir el-Zour, wouldn't have been possible without US intelligence.

The capture of Qaryatayn came as Syria and its strong backer Iran signed a joint memorandum of understanding for developing cooperation and coordination between the two countries' armies.

It said the memo was signed between the two countries' chiefs of staffs, adding that it provides for exchanging military expertise and intelligence and technology information in a way that can boost the two countries' capability for fight terrorism, according to state news agency SANA.

Iran has been one of Syrian President Bashar al-Assad's strongest supporters since the country's crisis began in 2011 and has sent thousands of Iranian-backed militiamen to boost his troops against opponents.

The Chief of StaffofIran's armed forces, Maj Gen Mohammad Bagheri, is on an official visit to Syria. He toured the front lines in the northern province of Aleppo and discussed military cooperation with the Syrian president.

Israel has been concerned about Iran's growing role in Syria and has been trying to keep the Islamic Republic and the fighters it backs away from its border.

Israel's Prime Minister Benjamin Netanyahu warned on Tuesday that he will not tolerate an Iranian military presence in neighbouring Syria.

Israel fears Iran will plant itself on the country's doorstep by establishing a Shia “corridor,” with land links from Iran to Lebanon, allowing the movement of fighters and weapons across the region.

Earlier on Saturday, the Syrian army said Israeli forces struck one of its positions near the Golan Heights south of the country.

The army statement said that Saturday's shelling by Israeli troops came after Syrian opposition fighters fired mortar rounds that hit an open area in the Israeli-occupied Golan Heights, giving the Jewish state a pretext to bomb the army.

The army said the shelling caused material damage without saying if there were casualties.

In one rare incident, Israel this week struck an anti-aircraft battery deep in Syria after it had opened fire on Israeli jets flying over Lebanon.

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